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An action is right for an individual if and only if it maximizes this individual's behavioural fitness.
Please note that this 'rightness criterion' does not mean that an individual's action would be right if and only if it maximizes this individual's behavioural fitness. A fitnessist who regularly eats other individuals in order to survive himself would not consider it to be right or good for himself to be eaten. Consequently fitnessism is 'non-universalizable', i.e. could not be held as right by everyone simultaneously without (ethical) disagreement being present, and also is indexical: Not only aesthetical propositions, but also ethical propositions are indexical in the same way as the word 'I' is indexical, which it is because what it denotes (i.e. to whom it refers) depends on who says it or has written it.
A person who for example expresses the proposition: "No individual ought ever to produce offspring." can never show that this strange opinion would be anything "higher", or even anything else, than this person's own personal opinion. Rather it is for this person's own part wrong for any individual to ever produce offspring. A predator about to catch a fleeing quarry shows through its behaviour that for its own part it would be right or good to eat the quarry. The quarry, on the other hand, shows through its behaviour that for its own part it would be wrong or bad to be eaten. Such are the ethical effects of natural selection.