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The line between copyediting and proofreading is narrow. Often, proof manuscripts contain typographical errors introduced during typesetting by the typesetter hitting incorrect keys (hence the word "typo" to refer to misplaced or incorrect characters). Earlier, when handwritten originals were common, it was often easier for a copyeditor to review and mark up a manuscript after it had been typeset and the proof copy produced. Today, proofreading usually refers to reviewing any text, hardcopy or electronic copy (on a computer), and checking for any kind of errors.
See also: copyediting